Date of Award

2008

Degree Type

Thesis

Degree Name

Master of Science

Program

Pharmacology and Toxicology

Supervisor

Dr. Stephen Ferguson

Second Advisor

Dr. Jane Rylett

Third Advisor

Dr. Susan Meakin

Abstract

Group I metabotropic glutamate receptors (mGluRs) - mGluRl and mGluR5 - are believed to contribute to neuronal death upon overstimulation and may be neuroprotective under other conditions. To develop treatment for mGluRs, we must identify proteins involved in their signalling. We have identified proline-rich tyrosine kinase 2 (Pyk2) as an mGluRl/5 interacting protein, and our objective was to investigate its contribution to the downstream activation of extracellular signal-regulated kinases (ERKs) by Group I mGluRs. In HEK 293 cells, ERK1/2 phosphorylation is greater in cells co-expressing mGluRl and wild-type Pyk2 than in cells expressing the receptor alone, and blocked in cells co-expressing the receptor and a dominant-negative mutant Pyk2. In cortical neurons, various inhibitors of endogenous Pyk2 block agonist stimulation of mGluRl-mediated ERK1/2 phosphorylation. Taken together, our data suggest wild-type Pyk2 is part of a cascade involved in enhancing ERK1∕2 activation by mGluRl.

Share

COinS
 
 

To view the content in your browser, please download Adobe Reader or, alternately,
you may Download the file to your hard drive.

NOTE: The latest versions of Adobe Reader do not support viewing PDF files within Firefox on Mac OS and if you are using a modern (Intel) Mac, there is no official plugin for viewing PDF files within the browser window.